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T??|5u? 010-62734371/62734373 5uP[?O?{ sui-office@cau.edu.cn suixiaolei@cau.edu.cn kXh??eg 2020t^12g02?e -N?V?QN'Yf[,g?yub?6R y??v T?y??tAAP?[?e?W?V?vKQ???S?R??xvzy??vw?bk?e?2021.03-2022.06y??v3u???N?`?Q?Y T-wSf~??]?S98019'`+RsY?Qu?eg1972.06.11gTf[?SNf[MOZS?XNN?b/gL??RYe?c?N?NNN,??f[y??v?i?Q .l/f iiru??T?S??@b?_??vCQ } ? iir???.l?vb__ g?e:g.l?T g:g.l ?]x`.l?T??`.l;N???~b?e:g.l ? ? g:g.l;N???v????e_/f(l?Wx?0(l?Wx?(W iir?vu?0?S???T?N"?ǐ z-N?S%c????v\O(u ? ?/f?V:N(l?Wx?/f iirSO?QT?yv??T?}v(?Tb?v?W,gCQ?N0 iir-N?e??/f??Q?/f???v(l?Wx???? ??????N蕄v蓽??}v?L?l?? ?sS(l?Wx?l???}v?Amino acid transporters ?AATs ?0 iirSO-N(l?Wx?l???}vR:N N'Y?[ ?e ?S?bATF/AAAP0APC?TMCF?[?e ??vMR:NbkxvzgY?v/fAAAP/ATF?[?e?vAAP?N?[?e?W?V0 (l?Wx???v??[?e(amino acid permease, AAP) ?/f?vMRxvzgY?v(l?Wx?l???}v?N?[?e0?bWS??AAP?N?[?e?W?V?vMRh??s?Q?v?^ir?T?~?~yr_'`??:_ ?N(W?R?? N ??N?[?ebXTKN??S g??'Y:S+R0?O?Y?AtAAP1;N???[MO(W(????MO ?(W?bWS???vP[?S0?sN?T9h-NGW gh??? ?;N???SN iir9h萄v(l?Wx?8T6eǐ z0AtAAP1??c?sN8T6e(l?Wx??[?PX[?}v(?Tb?T?yP[?N?w?0R?sQ????v\O(u?AtAAP2(W?bWS???g?[-Nh??? ?vQ?R???S??N(l?Wx??v???y????vsQ(Tegeder, 2014)0AtAAP3?SAtAAP5;N???N?[ iir?[?x'`(l?Wx??v8T6e ?vQ-NAtAAP3?v;N??h????MO:N9h?T??v痮v? ? ?AtAAP5d??N9h8T6e??S??N?yP[?S??-N?v(l?Wx???eQ?vsQ?dkY ?NAtAAP3 T?n'`?vя?v4l;zOsAAP3_N/fN*N?[MO?N(??? ??He?N?[V?(lx??T?|(lx?l???v?'`v?(Taylor et al., 2015)0AtAAP6x??N?T'`?v(l?Wx?l???}v ?;N???SN??c?bWS??[{?{RP[-N?v(l?Wx??v+T??SvQb?N(Tegeder, 2012)0xvz??ǐuir?Oo`f[?v?e?l?[?bWS??AAP?N?[?e?W?V?L??N?gr?SO?[MO ?fnx?NAtAAP?}v?v?~?gyr?_?SvQ?|?~?SsQ?| ?v^??:NAAP?N?[?ebXT?[ iir?v(l?Wx?l???T8T6e?N?S.l?N"?ǐ zw?0R?N?sQ????v\O(u(??I{ ?2016)0 ??t/fNLu??V^?l?y i?v????Vz?\OirKNN ??g?[/f????v?~Nm?^N?(uhV?[0(l?Wx?\O:N????v%?{QCQ } ?q_?T iir?vu??S?? ??[?S?N?X?R iir?S?~ }+T? ?OIQT\O(u?f?R?e?v ??[?X?RIQT\O(u?Nir?|?v+T??S?c?\Oir?T(??? g@w???\O(u ? ?,???S/f?NSO???S(l?Wx??v;N???ireg?nKNN ??Vdkxvz(l?Wx??v???ǐ z>f?v$\:N???0??HN(l?Wx?l???}v\O:N?N?[(l?Wx?蓽?l???v???}v ?(W(l?Wx??v???-Nw?@wsQ.??v\O(u ??cvzvQl???R??wQ g????v?[E?aIN0 ?S??e.s ?? ? RfSf ? Nggapg ? _? ? ?sSf?h ? ]?Us?. 2016. ?bWS??AAP?W?V?[?e?vuir?Oo`f[R?g?u}T?vSf[ ?36(3):372-378. Tegeder, M., 2014. Transporters involved in source to sink partitioning of amino acids and ureides: opportunities for crop improvement. Journal of Experimental Botany, 65(7): 1865-1878. Tegeder, M. Ward, J*,.24<>BHLVX\`bhl?????????ù????v?vhZL@?LZLhq%h?i)5?CJ PJhq%h ?5?>*CJ PJhq%h?~?5?>*CJ PJhq%hz?5?>*CJ PJhq%hGx/5?>*CJ PJhq%h?V5?6?>*CJ PJhq%h?V5?>*CJ PJhq%h?~?5?CJ PJhq%h ?5?CJ PJhq%h?i)5?CJ,hq%h ?5?CJ,hq%h ?5?CJ4PJhq%h?i)5?CJ4hq%h ?5?CJ4hq%h ?5?CJ4PJ hq%h ?.024b??< ? ? X f h j l ? ??????????????????$a$gd ?gd ??f???d??WD,]?f?`??gd?~????d??WD,]?`??gd?~? $?]?a$gd ?$ ?X ?]?a$gd ? $?r]?ra$gd ???????????? 0 6 8 : D F L T ~ ? ? ? ? ? ? ? ? ? ? ?  ??????????????|?pb?p|p?p??Thq%h?/?5?>*CJ PJh?1Yh?1Y5?CJ PJo(h?1Y5?>*CJ PJo(hS??5?>*CJ PJo(h?1Yh?1Y5?>*CJ PJo(h?1Y5?CJ PJo(hq%h??5?>*CJ PJh?5?>*CJ PJo(hq%h ?5?>*CJ PJhq%h ?5?CJ PJhq%h?~?5?>*CJ PJhq%h?O5?>*CJ PJhq%h?#?5?>*CJ PJ   & * , . 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FB?X ?????7???7?????d?0???????X ??? ????????? ????????? ????????? ????????4?4? a?f4yt|?N ^ d r x ? ? ?????? $$Ifa$? ? ? 6 5+??$IfWD?`??gd?@ & F$If?kd0$$If?4??B??FB?X ???? ?7?????d?0???????X ??????????????????????????????4?4? a?f4yt|?6 ? ??f??24 4P4????????i\ & F$Ifgd{?ckdC$$If???X X ?0???????X ??????????4?4? a? $7$8$H$Ifgd?????Z?$IfWD8?^??`?Z?gd?? $Ifgd] $Ifgd?@ ?4:@DPR`???????????????????????? ???????????????????xj?xj?xj?xj?xj?\h(cB*KHaJo(phh(cB*KH\?aJphh(cB*KH\?aJo(ph hq%h??B*KH\?aJphhq%h??5?B*\?aJphhq%h?w?5?B*\?aJphhq%h? ?aJhq%h?+aJhq%h>z?aJhq%h?w?aJhq%hDEpaJhq%h?X?aJhq%h?) aJhq%hDa?aJ$ <FHRf|v?????22?2?2?2?2?2?263?3?3?3?34 4444B4L4P4h4~4?4?4?????????????μ??????????????????xofhq%heu?aJhq%h?c?aJhq%hB%waJhq%h{?aJ hq%hxh? hq%hhZ?hq%h ?aJhq%hhZ?aJhq%h ?KHaJUhq%h??KHaJ hq%h??B*KH\?aJph hq%h??B*KH]?aJphh(cB*KHaJphhq%h??B*KHaJph(.M., 2012. Molecular Evolution of Plant AAP and LHT Amino Acid Transporters. Frontiers in Plant Science, 3(21): 1-11. Taylor, M.R., Reinders, A. Ward, J.M., 2015. Transport Function of Rice Amino Acid Permeases (AAPs). Plant Cell Physiology, 56(7): 1355-1363.?[?e??R?S?eHh(fnxT*N?s???[?W{Qf[uR?e???R?vwQSO`??S?8h?R?l) y??v?S?R?(WYe^?vc?[ N?ǐ?g??V?QY?vsQ?e.s?N? iir-N(l?Wx??v8T6eTb?MO ?(l?Wx?(W iir0W N?T0W N萄v????SRM??e_ ??N ?fnx iir-N(l?Wx?l???}v?v???'`06qT?ǐuir?Oo`f[?v?e_t??[?Q??t-N?vsQ?v(l?Wx?l???}v}?SOv^?L??W?V?vKQ???TRek?v?R?????? ??c?cRNA?v?c?S0?[?eg?IQ?[?PCR0u??k?R??????I{?vsQ?v?[???b/g ?gT?d?Q?~???bJT ??W{Qf[u?v?e.s??????R?T?yx?RKb???R ?wQSO?[?e?eHh?Y N? ?N ?y??v?[?ewQSO?eHh 1.y??vMRg?QY?]\O y??v?SN??g?я?Qt^?V?QY?vsQ?e.s ??N?,g???Wg?e?v?U\ ? 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