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Yu, Arabidopsis WRKY transcription factors WRKY12 and WRKY13 oppositely regulate flowering under short-day conditions[J]. Mol. Plant 9 (2016) 1492 1503. [7] H. Wang, et al., Overexpression of rice WRKY89 enhances ultraviolet B tolerance and disease resistance in rice plants[J]. Plant Mol. Biol. 65 (2007) 799 815. [8]F. Chen, et al., The WRKY transcription factor family in model plants and crops[J]. Crit. Rev.Plant Sci. 36 (2018) 1 25. [9] U. Phukan, G. Jeena, R. Shukla, WRKY transcription factors: molecular regulation and stress responses in plants[J]. Front. Plant Sci. 7 (2016) 760.?N0?[?e??R?S?eHh(fnxT*N?s???[?W{Qf[uR?e???R?vwQSO`??S?8h?R?l) 1.?e.s?g~bN??? (W?[^?vc?[ N ?y??v?S?R?????vsQ?e.s ??N?S_MRxvz?]\O?v?vsQ?s?r ??S?N?W{Qf[u?g??e.s?v???R0?[y??vZP?NEQR?N?T ??d?Q?[???eHh ??QS_ NG?0R?v? ??S?N;??pf[u?;N?d?Q?[???eHh?v???R ??W{Qf[u?[NN???W?vtQ??0 ?8h?e?l?f[u9hncy??vxvz?v?v??bN?N?[???eHh?S?b/g?~?N?~c?[Ye^ ?1u?^?8h?R0 ????eHh ?1 ??hKm?v??? 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